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Journal of Biotechnology 107 (2004) 193–232 Selectable marker genes in transgenic plants: applications, alternatives and biosafety Brian Miki, Sylvia McHugh Research Branch, Agriculture and Agri-Food Canada, Room 2091, KW Neatby Bldg., CEF, 960 Carling Avenue, Ottawa, Ont., Canada K1A 0C6 Received 18 July 2003; received in revised form 15 October 2003; accepted 27 October 2003